Unit VII: Genetics and Evolution · Chapter 4

Principles of Inheritance and Variation

Words to know

Word Simple meaning Example
Gene A unit of heredity that controls a trait Gene for plant height
Allele Different forms of the same gene T (tall) and t (dwarf)
Dominant The allele that shows up when paired with another allele T
Recessive The allele that is hidden when paired with a dominant allele t
Homozygous Both alleles are the same TT or tt
Heterozygous The two alleles are different Tt
Genotype The genes an organism has TT, Tt, tt
Phenotype How it looks Tall or dwarf

Why did Mendel choose peas?

  • Pea plants are easy to grow and have a short life cycle.
  • They have many clear, opposite traits (tall/dwarf, green/yellow).
  • They self-pollinate naturally but can also be cross-pollinated by hand.
  • Mendel studied 7 pairs of contrasting traits from 1856 to 1863.

Monohybrid cross: one trait at a time

Mendel crossed a pure tall (TT) plant with a pure dwarf (tt) plant.

Parents TT (tall) × tt (dwarf) → F1: all Tt (all tall) → self-pollinate Tt × Tt → F2: TT, Tt, Tt, tt → 3 tall : 1 dwarf (genotype 1 TT : 2 Tt : 1 tt).

Punnett square for F1 × F1 (Tt × Tt):

♀ \ ♂ T t
T TT (tall) Tt (tall)
t Tt (tall) tt (dwarf)

Test cross: find the hidden genotype

A tall plant could be TT or Tt. To find out, cross it with a dwarf (tt) plant:

  • If all the offspring are tall → the plant was TT.
  • If half are tall and half dwarf (1 : 1) → the plant was Tt.

Mendel's 3 laws

Law In simple words
1. Law of Dominance In a Tt plant, only the dominant trait (T, tall) shows
2. Law of Segregation The two alleles separate when gametes form, so each gamete gets only one allele. Also called the "law of purity of gametes"
3. Law of Independent Assortment Genes for different traits are passed on independently of each other

Dihybrid cross: two traits together

Mendel crossed round yellow (RRYY) seeds with wrinkled green (rryy) seeds. The F1 were all RrYy (round yellow). In the F2:

Round yellow Round green Wrinkled yellow Wrinkled green
9 3 3 1
  • 4 types of gametes from each parent: RY, Ry, rY, ry. That gives a 4 × 4 = 16 box Punnett square.
  • This cross proved the Law of Independent Assortment.

When Mendel's rules bend: exceptions

1. Incomplete dominance: blending

  • In the snapdragon (dog flower, Antirrhinum): red (RR) × white (rr) → pink (Rr) in F1.
  • F2 = 1 red : 2 pink : 1 white. Here the phenotype ratio is the same as the genotype ratio (1:2:1).

2. Co-dominance: both show

  • ABO blood groups in humans are controlled by the gene I, which has 3 alleles: Iᴬ, Iᴮ and i.
  • Iᴬ and Iᴮ are co-dominant (both show together as AB). i is recessive.
  • This also shows multiple alleles (more than 2 alleles for one gene in a population).
Genotype Blood group
IᴬIᴬ or Iᴬi A
IᴮIᴮ or Iᴮi B
IᴬIᴮ AB (co-dominance!)
ii O

3. Pleiotropy and polygenic inheritance

Pleiotropy Polygenic inheritance
Meaning One gene → many effects Many genes → one trait
Example Phenylketonuria Human skin colour, height

Chromosomal theory of inheritance

  • Sutton and Boveri said genes are on chromosomes, and chromosomes move in pairs like Mendel's factors.
  • Thomas Hunt Morgan proved it using fruit flies (Drosophila).
  • Linkage: genes on the same chromosome tend to be inherited together.
  • Recombination: new combinations appear when linked genes separate (by crossing over).
  • Tight linkage gives few recombinants; loose linkage gives more.
  • Alfred Sturtevant used recombination frequency to make the first gene maps.

Sex determination: boy or girl?

System Example Female Male
XX–XO Grasshopper XX XO (one X only)
XX–XY Humans, Drosophila XX XY
ZW–ZZ Birds ZW ZZ
Haplodiploidy Honey bee Diploid (32 chromosomes), from a fertilised egg Haploid (16 chromosomes), from an unfertilised egg

Mutation

  • A mutation is a change in DNA.
  • Point mutation: one base changes. Example: sickle-cell anaemia.
  • Frameshift mutation: a base is added or deleted, which shifts the whole reading frame.
  • Mutagens: things that cause mutation, such as UV rays and some chemicals.

Pedigree analysis

A pedigree is a family tree that shows how a trait is passed down. It helps track genetic diseases.

Symbol Meaning
Square Male
Circle Female
Shaded (filled) square or circle Affected person
Horizontal line joining a square and a circle Mating (marriage)
Double line Consanguineous mating (between relatives)

Genetic disorders

A. Mendelian disorders (a single gene change)

Disorder Type Key fact
Haemophilia X-linked recessive Blood doesn't clot. Mostly affects males; carrier mothers pass it to sons. Famous in Queen Victoria's family
Colour blindness X-linked recessive Can't tell red from green. About 8% of males and 0.4% of females
Sickle-cell anaemia Autosomal recessive Glutamic acid → valine at the 6th position of the β-globin chain (GAG → GUG). RBCs become sickle-shaped
Phenylketonuria Autosomal recessive Can't convert phenylalanine → tyrosine, which leads to mental retardation
Thalassemia Autosomal recessive Less globin is made (a quantity problem). Sickle-cell makes the wrong globin (a quality problem)

B. Chromosomal disorders (whole chromosome changes)

  • Aneuploidy: one chromosome extra or missing, caused by failed separation during cell division.
  • Polyploidy: whole extra sets of chromosomes. Common in plants.
Disorder Chromosomes Features
Down's syndrome Trisomy 21 (47 chromosomes) Short, small round head, furrowed tongue, partly open mouth, palm crease, slow mental development
Klinefelter's syndrome 47, XXY Male with some female features (breast growth); sterile
Turner's syndrome 45, XO Female with underdeveloped ovaries; sterile

30-second revision

  • Monohybrid 3:1 (phenotype), 1:2:1 (genotype); dihybrid 9:3:3:1; test cross 1:1.
  • Incomplete dominance: snapdragon → pink (1:2:1).
  • ABO blood groups = co-dominance + multiple alleles (3-6-4).
  • Morgan → Drosophila → linkage; Sturtevant → gene maps.
  • Birds: female ZW. Honey bee male: haploid (16).
  • Haemophilia and colour blindness: X-linked recessive. Sickle-cell: Glu6Val.
  • Down's = 21 trisomy; Klinefelter = XXY; Turner = XO.